For any given gene, what ultimately determines which dna strand serves as the template strand?

The answer is the base sequence of gene's promoter.

DNA is a two-stranded helix. One strand is called a template strand, the other one is called a coding strand. The template strand is copied into mRNA, and the coding strand is complementary to the template strand. RNA polymerase, an enzyme responsible for transcription, has to recognize the right strand before transcription begins. So, it recognizes one or more short sequences on the coding strand. This is the base sequence of gene's promoter. Thus, the complementary strand is the template strand.

Ifound this on google “concept 13: meiosis ii: telophase ii. a nuclear envelope forms around each set of chromosomes and cytokinesis occurs, producing four daughter cells, each with a haploid set of chromosomes. a nuclear envelope forms around each set of chromosomes.” i hope this you for what your looking for.